Skipper, Kowalski, Rico and Private are an elite strike force with top-tier commando skills and a world-class secret headquarters. At least, that's how Skipper would describe them. Others might say they're four cute penguins living among their animal brethren at the Central Park Zoo. But no matter, the penguins are too busy maintaining order to care what others think.
Only one thing is capable of distracting them from their mission: their obnoxious new neighbour, King Julien. This party animal, and self-proclaimed 'King of the Lemurs,' has just arrived at the zoo and he's looking to shake things up a bit. Uh oh, there goes the neighbourhood!